Royal Mail employees and driving examiners are also striking.
Snow, ice, and fog impeded road and air travel on Tuesday, when rail workers walked out.
On Wednesday, half of rail lines will be closed, and most of Scotland and Wales will have no service.
This week could potentially see the first-ever nationwide nurses' strike.
Friday, rail, bus, baggage, highway, and driving examiners will strike.
Later this week, the government's emergency Cobra committee will meet again to discuss industrial action.
115,000 CWU members strike during Royal Mail's busiest time of year, when people and businesses send Christmas cards and gifts.
Some parcel companies say the Royal Mail strike is delaying next-day deliveries as customers look for alternatives.
DPD Group said, "We are experiencing short delays to our next-day delivery service in a few places due to Royal Mail's industrial action, which has affected the entire sector."
Evri, previously Hermes, said harsh weather, Royal Mail strikes, and staff shortages cause "localized delays."
Yodel claimed some delivery times are longer, but didn't say why.
Royal Mail employees will also strike on December 23 and Christmas Eve.
The conflict has been going on since the summer, and money is a crucial issue.
As costs grow, workers want raises. Inflation is over 11%, the fastest pace in over 40 years.
Food and energy prices have risen.
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